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Most of the penis develops from the same embryonic tissue as does the clitoris in females. The skin around the penis and the urethra come from the same embryonic tissue from which develops the labia minora in females.

Erections can also occur in non-sexual situations; spontaneous non-sexual erections frequently occur during adolescence and during sleep.

In its flaccid unerect state, the shaft of the penis has the feel of a dense sponge encased in very smooth eyelid-type skin. The glans of the penis, in uncircumcised males, is covered by the foreskin.

In its fully erect hard state, the shaft of the penis is rigid, with the skin tightly stretched. The glans of the erect penis has the feel of a raw mushroom.

An erect penis may be straight or curved and may point at an upward angle, a downward angle, or straight ahead. As of [update] , the best research on penis size concluded that the average erect human penis is The most common form of alteration of the penis is circumcision , which is the removal of part or all of the foreskin for various cultural, religious, and more rarely medical reasons.

There is controversy surrounding circumcision. The human penis is made up of three columns of tissue : two corpora cavernosa lie next to each other on the dorsal side and one corpus spongiosum lies between them on the ventral side.

The enlarged and bulbous-shaped end of the corpus spongiosum forms the glans penis with two specific types of sinusoids, which supports the foreskin , or prepuce, a loose fold of skin that in adults can retract to expose the glans.

The rounded base of the glans is called the corona. The perineal raphe is the noticeable line along the underside of the penis.

It is a passage both for urine and for the ejaculation of semen. Sperm are produced in the testes and stored in the attached epididymis.

During ejaculation, sperm are propelled up the vas deferens , two ducts that pass over and behind the bladder.

Fluids are added by the seminal vesicles and the vas deferens turns into the ejaculatory ducts , which join the urethra inside the prostate gland.

The prostate as well as the bulbourethral glands add further secretions, and the semen is expelled through the penis. The raphe is the visible ridge between the lateral halves of the penis, found on the ventral or underside of the penis, running from the meatus opening of the urethra across the scrotum to the perineum area between scrotum and anus.

The human penis differs from those of most other mammals , as it has no baculum or erectile bone and instead relies entirely on engorgement with blood to reach its erect state.

A distal ligament buttresses the glans penis and plays an integral role to the penile fibroskeleton, and the structure is called "os analog," a term coined by Geng Long Hsu in the Encyclopedia of Reproduction.

The human penis cannot be withdrawn into the groin, and it is larger than average in the animal kingdom in proportion to body mass. Penile measurements vary, with studies that rely on self-measurement reporting a significantly higher average size than those which rely on measurements taken by health professionals.

As of [update] , a systematic review of 15, men and the best research to date on the topic, as the subjects were measured by health professionals concluded that the average length of an erect human penis is Among all primates, the human penis is the largest in girth, but is comparable to the chimpanzee penis and the penises of certain other primates in length.

It was In the developing fetus, the genital tubercle develops into the glans of the penis in males and into the clitoral glans in females; they are homologous.

The urogenital fold develops into the skin around the shaft of the penis and the urethra in males and into the labia minora in females.

On entering puberty , the penis, scrotum and testicles will enlarge toward maturity. During the process, pubic hair grows above and around the penis.

A large-scale study assessing penis size in thousands of to year-old males found no difference in average penis size between year-olds and year-olds.

From this, it can be concluded that penile growth is typically complete not later than age 17, and possibly earlier. In males, the expulsion of urine from the body is done through the penis.

The urethra drains the bladder through the prostate gland where it is joined by the ejaculatory duct , and then onward to the penis.

At the root of the penis the proximal end of the corpus spongiosum lies the external sphincter muscle.

This is a small sphincter of striated muscle tissue and is in healthy males under voluntary control. Relaxing the urethra sphincter allows the urine in the upper urethra to enter the penis properly and thus empty the urinary bladder.

Physiologically, urination involves coordination between the central , autonomic , and somatic nervous systems. In infants, some elderly individuals, and those with neurological injury, urination may occur as an involuntary reflex.

Brain centers that regulate urination include the pontine micturition center , periaqueductal gray , and the cerebral cortex.

The distal section of the urethra allows a human male to direct the stream of urine by holding the penis. This flexibility allows the male to choose the posture in which to urinate.

In cultures where more than a minimum of clothing is worn, the penis allows the male to urinate while standing without removing much of the clothing.

It is customary for some men to urinate in seated or crouched positions. The preferred position may be influenced by cultural or religious beliefs.

A meta-analysis [24] summarizing the evidence found no superior position for young, healthy males.

For elderly males with LUTS however, in the sitting position compared to the standing:. This urodynamic profile is related to a lower risk of urologic complications, such as cystitis and bladder stones.

An erection is the stiffening and rising of the penis, which occurs during sexual arousal , though it can also happen in non-sexual situations.

Spontaneous erections frequently occur during adolescence due to friction with clothing, a full bladder or large intestine, hormone fluctuations, nervousness, and undressing in a nonsexual situation.

It is also normal for erections to occur during sleep and upon waking. See nocturnal penile tumescence.

The primary physiological mechanism that brings about erection is the autonomic dilation of arteries supplying blood to the penis, which allows more blood to fill the three spongy erectile tissue chambers in the penis, causing it to lengthen and stiffen.

The now-engorged erectile tissue presses against and constricts the veins that carry blood away from the penis. More blood enters than leaves the penis until an equilibrium is reached where an equal volume of blood flows into the dilated arteries and out of the constricted veins; a constant erectile size is achieved at this equilibrium.

The scrotum will usually tighten during erection. Erection facilitates sexual intercourse though it is not essential for various other sexual activities.

Although many erect penises point upwards see illustration , it is common and normal for the erect penis to point nearly vertically upwards or nearly vertically downwards or even horizontally straight forward, all depending on the tension of the suspensory ligament that holds it in position.

The following table shows how common various erection angles are for a standing male, out of a sample of 1, males aged 20 through In the table, zero degrees is pointing straight up against the abdomen, 90 degrees is horizontal and pointing straight forward, while degrees would be pointing straight down to the feet.

An upward pointing angle is most common. Ejaculation is the ejecting of semen from the penis, and is usually accompanied by orgasm.

A series of muscular contractions delivers semen, containing male gametes known as sperm cells or spermatozoa , from the penis. It is usually the result of sexual stimulation.

Rarely, it is due to prostatic disease. Ejaculation may occur spontaneously during sleep known as a nocturnal emission or wet dream.

Anejaculation is the condition of being unable to ejaculate. Ejaculation has two phases: emission and ejaculation proper.

The emission phase of the ejaculatory reflex is under control of the sympathetic nervous system , while the ejaculatory phase is under control of a spinal reflex at the level of the spinal nerves S2—4 via the pudendal nerve.

A refractory period succeeds the ejaculation, and sexual stimulation precedes it. The human penis has been argued to have several evolutionary adaptations.

The purpose of these adaptations is to maximise reproductive success and minimise sperm competition. Sperm competition is where the sperm of two males simultaneously resides within the reproductive tract of a female and they compete to fertilise the egg.

This is the process whereby males unwittingly invest their resources into offspring of another male and, evolutionarily speaking, should be avoided.

The most researched human penis adaptations are testis and penis size , ejaculate adjustment and semen displacement.

Evolution has caused sexually selected adaptations to occur in penis and testis size in order to maximise reproductive success and minimise sperm competition.

Sperm competition has caused the human penis to evolve in length and size for sperm retention and displacement. As a result, this adaptation also leaves the sperm less vulnerable to sperm displacement and semen loss.

Another reason for this adaptation is that, due to the nature of the human posture, gravity creates vulnerability for semen loss. Therefore, a long penis, which places the ejaculate deep in the vaginal tract, could reduce the loss of semen.

Another evolutionary theory of penis size is female mate choice and its associations with social judgements in modern-day society. These varied in height, body shape and flaccid penis size, with these aspects being examples of masculinity.

This is reflected in the association between believed sexual prowess and penis size and the social judgement of penis size in relation to 'manhood'.

Like the penis, sperm competition has caused the human testicles to evolve in size through sexual selection. The human testicles are moderately sized when compared to other animals such as gorillas and chimpanzees, placing somewhere midway.

Research has also demonstrated that evolutionary adaptations of testis size are dependent on the breeding system in which the species resides.

Human males live largely in monogamous societies like gorillas, and therefore testis size is smaller in comparison to primates in multi-male breeding systems, such as chimpanzees.

The reason for the differentiation in testis size is that in order to succeed reproductively in a multi-male breeding system, males must possess the ability to produce several fully fertilising ejaculations one after another.

One of the primary ways in which a male's ejaculate has evolved to overcome sperm competition is through the speed at which it travels. Ejaculates can travel up to 30—60 centimetres at a time which, when combined with its placement at the highest point of the vaginal tract, acts to increase a male's chances that an egg will be fertilised by his sperm as opposed to a potential rival male's sperm , thus maximising his paternal certainty.

In addition, males can—and do—adjust their ejaculates in response to sperm competition and according to the likely cost-benefits of mating with a particular female.

The number of sperm in any given ejaculate varies from one ejaculate to another. A male will alter the number of sperm he inseminates into a female according to his perceived level of sperm competition, [29] inseminating a higher number of sperm if he suspects a greater level of competition from other males.

In support of ejaculate adjustment, research has shown that a male typically increases the amount he inseminates sperm into his partner after they have been separated for a period of time.

Increasing the number of sperm a male inseminates into a female acts to get rid of any rival male's sperm that may be stored within the female, as a result of her potential extra-pair copulations EPCs during this separation.

Through increasing the amount he inseminates his partner following separation, a male increases his chances of paternal certainty.

This increase in the number of sperm a male produces in response to sperm competition is not observed for masturbatory ejaculates. Males also adjust their ejaculates in response to sperm competition in terms of quality.

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Erect human penis. Normal erect penis. Downward inclined erect penis. Frontal view of erect, vertically inclined, uncircumcised penis.

X1 Frontal view of glans. X2Side view of glans. Penis circoncis en erection. Oriental male with erection.

Normal 4. Normal erection with slight upward inclination. Erect human penis with shaved genitalia. Asian male. Non-retracted and retracted foreskin of an erect penis.

Normal upward inclined erection. Circumcised penis with full perpendicular erection. Circumcised penis, flacid and erect.

Normal erect penis: model thrusting buttocks forward. A normal erect human penis with detail of glans. Normal erect penis viewed from side and above.

Erect penis with foreskin removed. Normal erect penis under different lighting or exposure. Normal erect penis with upward inclination. Normal uncircumcised human penis erect.

Glans penis left: flaccid penis; right: erect penis. Corona of the glans penis erect penis. Glans penis with frenulum left: flaccid penis; right: erect penis.

Penises can become distorted by disorders, injury or repeated mild maltreatment, but deformation rarely affects function.

Upward curve: possibly the result of a preference for pressing vigourously against a hard surface.

Prince Albert Glans and Frenulum piercing. Tatoo, constricting ring and frenulum piercing. There are many images and videos of masturbation within Wikimedia Commons, of which these are only illustrative samples:.

Circumcised man masturbating using fist grip. Radically circumcised Male masturbation using light, low impact two finger technique.

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